Staab M, Achury R, Ammer C, Ehbrecht M, Irmscher V, Mohr H, Schall P, Weisser WW, Blüthgen N (2022) Negative effects of forest gaps on dung removal in a full-factorial experiment. Journal of Animal Ecology, accepted. Link

Liu X, Huang Y, Chen L, Li S, Bongers FJ, Castro-Izaguirre N, Liang Y, Yang B, Chen Y, Schnabel F, Tang T, Xue Y, Trogisch S, Staab M, Bruelheide H, Schmid B, Ma K (2022) Species richness, functional traits and climate interactively affect tree survival in a large forest biodiversity experiment. Journal of Ecology, accepted. Link

Rappa NJ, Staab M, Frey J, Winiger N, Klein AM (2022) Multiple forest structural elements are needed to promote beetle biomass, diversity and abundance.  Forest Ecosystems 9: 100056. Link

Blüthgen N, Staab M, Achury R, Weisser WW (2022) Unravelling insect declines: can space replace time? Biology Letters 18: 20210666. Link

Hoenle PO, Donoso DA, Argoti A, Staab M, von Beeren C, Blüthgen N (2022) Rapid ant community re-assembly in a Neotropical forest: recovery dynamics and land-use legacy. Ecological Applications 32: e2559. Link

Pedersen KM, Blüthgen N (2022) Seed size and pubescence facilitate secondary dispersal by dung beetles. Biotropica DOI: 10.1111/btp.13052

Ceron K, Diogo B. Provete DB, Pires MM, Araujo AC, Blüthgen N, Santana DJ (2022) Prey availability and interaction rewiring drive the spatial and seasonal structure of a predator-prey metaweb. Ecology

Melo Teles Gomes IJ, Solar R, Blüthgen N, Vasconcelos HL, Basilio Silva LE, Oliveira KE, Moreira MZ, Campos RI (2022) Multiple effects of mutualistic ants improve the performance of a neotropical ant-plant: A long-term study with the Cecropia-Azteca system. Basic and Applied Ecology 57: 78-90

Scherreiks P, Gossner MM, Ambarlı D., Ayasse M, Blüthgen N, Fischer M, Klaus VH, Kleinebecker T, Neff F, Prati D, Seibold S, Simons NK, Weisser WW, Wells K, Westphal C, Thiele J (2022) Present and historical landscape structure shapes current species richness in Central European grasslands. Landscape Ecology DOI: 10.1007/s10980-021-01392-7


​McKenzie S.K., Winston, M.E., Grewe F., Vargas Asensio G., Rodríguez-Hernández N., Rubin B.E.R., Murillo-Cruz C., von Beeren C., Moreau C.S., Suen G., Pinto-Tomás A.A., & Kronauer D.J.C. (2021). The genomic basis of army ant chemosensory adaptations. Molecular Ecology,

von Beeren C, Brückner  A, Hoenle  PO, Ospina-Jara B, Kronauer DJC, & Blüthgen N. (2021). Multiple phenotypic traits as triggers of host attacks towards ant symbionts: body size, morphological gestalt, and chemical mimicry accuracy. Frontiers in Zoology, 18:46;

von Beeren C, Blüthgen  N, Hoenle  PO, Pohl  S, Brückner  A, Tishechkin  AK, Maruyama  M, Brown  BV, Hash  JM, Hall  WE, Kronauer DJC (2021) A remarkable legion of guests: Diversity and host specificity of army ant symbionts. Molecular Ecology, DOI: 10.1111/mec.16101

Blüthgen N, Staab M (2021) Ecology: Mammals, interactions networks and the relevance of scale. Current Biology 31: R850-R853. Link

Fornoff F, Staab M, Zhu CD, Klein AM (2021) Multi-trophic communities re-establish with canopy cover and microclimate in a subtropical forest biodiversity experiment. Oecologia 196: 289-301. Link

Zumstein P, Bruelheide H, Fichtner A, Schuldt A, Staab M, Härdtle W, Zhao H, Assmann T (2021) What shapes ground beetle assemblages in a tree species-rich subtropical forest? ZooKeys 1044: 907-927. Link

Guo P, Wang M, Orr M, Li Y, Chen J, Zhou Q, Staab M, Fornoff F, Chen G, Zhang N, Klein AM, Zhu CD (2021) Tree diversity promotes predatory wasps and parasitoids but not pollinator bees in a subtropical experimental forest. Basic and Applied Ecology 53: 134-142. Link

Wehner, K., R. Schuster, N.K. Simons, R.A. Norton, N. Blüthgen & M. Heethoff (2021): How land-use intensity affects sexual and parthenogenetic oribatid mites in temperate forests and grasslands in Germany. Experimental & Applied Acarology (online first). doi:10.1007/s10493-020-00586-z Link

Wu D, Staab M, Yu M (2021) Canopy closure retards fine wood decomposition in subtropical regenerating forests. Ecosystems 24: 1875-1890. Link

Wu D, Pietsch KA, Staab M, Yu M (2021) Wood species identity alters dominant factors driving fine wood decomposition along a tree diversity gradient in subtropical plantation forests. Biotropica 53: 643-657. Link

Staab M, Liu X, Assmann T, Bruelheide H, Buscot F, Durka W, Erfmeier A, Klein AM, Ma K, Michalski S, Wubet T, Schmid B, Schuldt A (2020) Tree phylogenetic diversity structures multitrophic communities. Functional Ecology 35: 521-534. Link

Wehner K, Renker C, Simons NK, Weisser WW, Blüthgen N (2021): Narrow environmental niches predict land-use responses and vulnerability of land snail assemblages. BMC Ecology & Evolution 21(15). doi:10.1186/s12862-020-01741-1 Link

Simons NK, Felipe-Lucia MR, Schall P, Ammer C, Bauhus J, Blüthgen N, Boch S, Buscot F, Fischer M, Goldmann K, Gossner MM, Hänsel F, Jung K, Manning P, Nauss T, Oelmann Y, Pena R, Polle A, Renner SC, Schloter M, Schöning I, Schulze ED, Solly EF, Sorkau E, Stempfhuber B, Wubet T, Müller J, Seibold S, Weisser WW (2021) National Forest Inventories capture the multifunctionality of managed forests in Germany. Forest Ecosystems 8(5). doi:10.1186/s40663-021-00280-5 Link

Seibold S, Hothorn T, Gossner MM, Simons NK, Blüthgen N, Müller J, Ambarlı D, Ammer C, Bauhus J, Fischer M, Habel JC, Penone C, Schall P, Schulze ED, Weisser WW (2021) ​Insights from regional and short‐term biodiversity monitoring datasets are valuable: a reply to Daskalova et al. 2021. Insect Conservation and Diversity 14: 144-148 Link

CaraDonna P, Burkle L, Schwarz B, Resasco J, Knight T, Benadi G, Blüthgen N, Dormann C, Fang Q, Fründ J, Gauzens B, Kaiser-Bunbury C, Winfree R, Vazquez D (2021) Seeing through the static: The temporal dimension of plant–animal mutualistic interactions. Ecology Letters 24: 149-161

Thiel T, Gaschler S, Mody K, Blüthgen N, Drossel B (2020) Impact of herbivore preference on the benefit of plant trait variability. Theoretical Ecology 14: 173-187


Neumüller U, Burger H, Krausch S, Blüthgen N, Ayasse M (2020) Interactions of local habitat type, landscape composition and flower availability moderate wild bee communities. Landscape Ecology 35: 2209-2224

Ambarlı D, Simons NK, Wehner K, Kämper W, Gossner MM, Nauss T, Neff F, Seibold S, Weisser WW, Blüthgen N (2020) ​Animal-mediated ecosystem process rates in forests and grasslands are affected by climatic conditions and land-use intensity. Ecosystems 24: 467-483 Link

Hoenle PO, Lattke JE,  Donoso DA, von Beeren CHeethoff MSchmelzle S, Argoti A, Camacho L, Stroebel BBluethgen N (2020)  Odontomachus davidsoni sp. nov. (Hymenoptera, Formicidae), a new conspicuous trap-jaw ant from Ecuador. Zookeys 948: 75-105

Mody K, Lerch D, Müller AK, Simons NK, Blüthgen N, Harnisch M (2020) Flower power in the city: Replacing roadside shrubs by wildflower meadows increases insect numbers and reduces maintenance costs. PLOS ONE 15(6): e0234327. doi:10.1371/journal.pone.0234327

Thiel T, Gaschler S, Mody K, Blüthgen N, Drossel B (2020) Impact of plant defense level variability on specialist and generalist herbivores. Theoretical Ecology 13: 409-424

Schwarz B, Vázquez DP, CaraDonna PJ, Knight TM, Benadi G, Dormann CF, Gauzens B, Motivans E, Resasco J, Blüthgen N, Burkle LA, Fang Q, Kaiser–Bunbury CN, Alarcón R, Bain JA, Chacoff NP, Huang SQ, LeBuhn G, MacLeod M, Petanidou T, Rasmussen C,  Simanonok MP, Thompson AH, Fründ J (2020) Temporal scale‐dependence of plant–pollinator networks. Oikos 129: 1289-1302

Wehner K, Schäfer L, Blüthgen N, Mody K (2020) Seed type, habitat and time of day influence post-dispersal seed removal in temperate ecosystems. PeerJ 8: e8769


Seibold, S., Gossner, M.M., Simons, N.K., Blüthgen, N., Müller, J., Ambarli, D., Ammer, C., Bauhus, J., Fischer, M., Habel, J.C., Linsenmair, K.E., Nauss, T., Penone, C., Prati, D., Schall, P., Schulze, E.-D., Vogt, J., Wöllauer, S. & Weisser W.W. (2019): Arthropod decline in grasslands and forests is associated with landscape-level drivers. Nature. 574:671–674.

Vogt, J., Klaus V.H., Both S., Fürstenau C., Gockel S., Gossner M.M., Heinze J., Hemp A., Hölzel N., Jung K., Till K., Lauterbach R., Lorenzen K., Ostrowski A., Otto N., Prati D., Renner S., Schumacher U., Seibold S., Simons N.K., Steitz I., Teuscher M., Thiele J., Weithmann S., Wells K., Wiesner K., Ayasse M., Blüthgen N., Fischer M., Weisser W.W. (2019). Eleven years’ data of grassland management data in Germany. Biodiversity Data Journal. 7: e36387

Neff F., Blüthgen N., Chisté, M.N.Simons N.K., Steckel J., Weisser W.W., Westphal C., Pellissier, L., Gossner M.M. (2019). Cross-scale effects of land use on the functional composition of herbivorous insect communities. Landscape Ecology 34: 2001-2015

Wehner K., Renker C., Brückner A., Simons N.K., Weisser W.W., Blüthgen N. (2019): Land‐use in Europe affects land snail assemblages directly and indirectly by modulating abiotic and biotic drivers, Ecosphere 10: e02726

Heinze J., Simons N.K., Seibold S., Wacker A., Weithoff G., Gossner M.M., Prati D., Bezemer T.M., Joshi J. (2019). The relative importance of plant-soil feedbacks for plant-species performance increases with decreasing intensity of herbivory, Oecologia. 190(3):651–664.

Lizon à l'Allemand S., Brückner A., Hashim R., Witte V.*, von Beeren C.* (2019). Competition as possible driver of dietary specialisation in the mushroom harvesting ant Euprenolepis procera (Hymenoptera: Formicidae).  Myrmecological News, 29: 71-91 *contributed equally

Hoenle P., Blüthgen N., Brückner A., Kronauer D.J.C., Fiala B., Donoso D., Smith M.A., Ospina Jara B., von Beeren C. (2019).  Species-level predation network uncovers high prey specificity in a Neotropical army ant community. Molecular Ecology, 28: 2423-2440​


Ekroos J, Kleijn D, Batáry P, Albrecht M, Báldi A, Blüthgen N, Knop E, Kovács-Hostyánszki A, Smith HG (2019) High land-use intensity in grasslands constrains wild bee species richness in Europe. Biological Conservation 241: 108255

Camara T, Leal IR, Oliveira F, Blüthgen N, Arnan X (2019) Anthropogenic disturbance and rainfall variation threaten the stability of plant-ant interactions in the Brazilian Caatinga. Ecography 42: 1960-1972


Arnan X, Molowny-Horas R, Blüthgen N (2019) Food resource exploitation and functional resilience in ant communities found in common Mediterranean habitats. Science of the Total Environment 684: 126-135

Schmelzle S, Blüthgen N (2019) Under pressure: force resistance measurements in box mites (Actinotrichida, Oribatida). Frontiers in Zoology 16: 24

Busch V, Klaus VH, Schaefer D, Prati D, Boch S, Müller J, Socher SA, Chisté M, Mody K, Blüthgen N, Fischer M, Hölzel N, Kleinebecker T (2019) Will I stay or will I go? Plant species specific response and tolerance to high land-use intensity in temperate grassland ecosystems. Journal of Vegetation Science 30: 674-686


Happe AK, Alins G, Blüthgen N, Boreux V, Bosch J, García D, Hambäck PA, Klein AM, Martínez-Sastre R, Miñarro M, Müller AK, Porcel M, Rodrigo A, Roquer-Beni L, Samnegård U, Tasin M, Mody K (2019) Predatory arthropods in apple orchards across Europe: Responses to agricultural management, adjacent habitat, landscape composition and country. Agriculture Ecosystems and Environment 273: 141-150


Fornoff F, Klein AM, Blüthgen N, Staab M (2019) Tree diversity increases robustness of multi-trophic interactions. Proceedings of the Royal Society B 286: 2399


Brückner A, Hoenle P, von Beeren C (2018). Comparative chemical analysis of army ant mandibular gland volatiles (Formicidae: Dorylinae). PeerJ  6:e5319

O ’ Donnell S, Bulova S, Barrett M,  von Beeren C (2018). Brain investment under colony-level selection: soldier specialization in Eciton army ants (Formicidae: Dorylinae). BMC Zoology, 3:3

Frank K, Krell FT, Walter P, Slade EM, Raine EH, Chiew LY, Vairappan CS, Schmitt T, Blüthgen N (2018) Global dung webs: high trophic generalism of dung beetles along the latitudinal diversity gradient. Ecology Letters 21: 1229-1236

Wehner, K., Heethoff, M., Brückner, A. (2018). Seasonal fluctuation of oribatid mite communities in forest microhabitats. PeerJ 6: e4863.

Heethoff, M. (2018). Cryptic species - conceptual or terminological chaos? Trends in Ecology and Evolution 33: 310.

Ströbel, B., Schmelzle, S., Blüthgen, N., Heethoff, M. (2018). An automated device for the digitization and 3D modeling of insects, combining extended-depth-of-field and all-side multi-view imaging. ZooKeys 759: 1-27.

Tschulakow AV, Oltrup T, Bende T, Schmelzle S, Schraermeyer​ U (2018) The anatomy of the foveola reinvestigated. PeerJ 6: e4482

Brückner A, Schuster R, Smit T, Heethoff M (2018) Imprinted or innate food preferences in the model mite Archegozetes longisetosus (Actinotrichida, Oribatida, Trhypochthoniidae). Soil Organisms 90: 23-26

Wehner K, Heethoff M, Brückner A (2018) Sex ratios of oribatid mites differ among microhabitats. Soil Organisms 90: 13-22


Brückner A, Schuster R, Wehner K, Heethoff M (2018) Effects of nutritional quality on the reproductive biology of Archegozetes longisetosus (Actinotrichida, Oribatida, Trhypochthoniidae). Soil Organisms 90: 1-12

von Beeren C, Brückner A, Maruyama M, Burke G, Wieschollek J, Kronauer JC (2018). Chemical and behavioral integration of army ant-associated rove beetles -  a comparison between specialists and generalists. Frontiers in Zoology 15:8.

Brückner A, Heethoff M (2018) Nutritional effects on chemical defense alter predator-prey dynamics. Chemoecology 28: 1-9.

Bergmann P, Laumann M, Norton RA, Heethoff M (2018) Cytological evidence for automictic thelytoky in parthenogenetic oribatid mites (Acari, Oribatida): Synaptonemal complexes confirm meiosis in Archegozetes longisetosus. Acarologia 58: 342-356.

Brückner A, Schuster R, Smit T, Pollierer MM, Schäffler I, Heethoff M (2018) Track the snack - olfactory cues shape foraging behaviour of decomposing soil mites (Oribatida). Pedobiologia 66: 74-80

Happe A-K, Riesch F, Rösch V, Gallé R, Tscharntke T & Batáry P (2018) Small-scale agricultural landscapes and organic management support wild bee communities of cereal field boundaries. Agriculture, Ecosystems and Environment 254: 92-98

Chisté MN, Mody K, Kunz G, Gunczy J, Blüthgen N (2018) Intensive land use drives small-scale homogenization of plant- and leafhopper communities and promotes generalists. Oecologia 186: 529-540

Rosumek FB, Blüthgen N, Brückner A, Menzel F, Gebauer G, Heethoff M (2018). Unveiling community patterns and trophic niches of tropical and temperate ants using an integrative framework of field data, stable isotopes and fatty acids. PeerJ 6: e5467


Felipe-Lucia MR, Soliveres S, Penone C, Manning P, van der Plas F, Boch S, Prati D, Ammer C, Schall P, Gossner MM, Bauhus J, Buscot F, Blaser S, Blüthgen N, de Frutos A, Ehbrecht M, Frank K, Goldmann K, Hänsel F, Jung K, Kahl T, Nauss T, Oelmann Y, Pena R, Polle A, Renner S, Schloter M, Schöning I, Schrumpf M, Schulze ED, Solly E, Sorkau E, Stempfhuber B, Tschapka M, Weisser WW, Wubet T, Fischer M, Allan E (2018) Multiple forest attributes underpin the supply of multiple ecosystem services. Nature Communications 9: 4839

Mesquita-Neto J, Blüthgen N, Schlindwein C (2018) Flowers with poricidal anthers and their complex interaction networks - disentangling legitimate pollinators and illegitimate visitors. Functional Ecology 32: 2321-2332


van der Plas F, Allan E, Fischer M, Alt F, Arndt H, Binkenstein J, Blaser S, Blüthgen N, Boehm S, Hoelzel N, Klaus V, Kleinebecker T, Morris K, Oelmann Y, Prati D, Renner S, Rillig MC, Schaefer HM, Schloter M, Schmitt B, Schoening I, Schrumpf M, Solly E, Sorkau E, Steckel J, Steffan-Dewenter I, Stempfhuber B, Tschapka M, Weiner C, Weisser W, Werner M, Westphal C, Wilcke W, Manning P (2018) Towards the development of general rules describing landscape heterogeneity-multifunctionality relationships. Journal of Applied Ecology 56: 168-179


Costa F, Blüthgen N, Viana-Junior A, Guerra TJ, Di Spirito L, Neves FS (2018) Resilience to fire and climate seasonality drive the temporal dynamics of ant-plant interactions in a fire-prone ecosystem. Ecological Indicators 93: 247-255


Frank K, Brückner A, Blüthgen N, Schmitt T (2018) In search for cues: dung beetle attraction and the significance of volatile composition in dung. Chemoecology 28: 145-152


Câmara T, Leal I, Blüthgen N, Oliveira FM, de Queiroz R, Arnan X (2018) Effects of chronic anthropogenic disturbance and rainfall on the specialization of ant-plant mutualistic networks in the Caatinga, a Brazilian dry forest. Journal of Animal Ecology 87: 1022-1033


Lösel P, Heuveline V, Jerome NT, Kopmann A, van de Kamp T, Schmelzle S, Heethoff M (2017) Biomedisa: The Biomedical Image Segmentation App. 3rd International Conference on Tomography of Materials and Structures Lund, Sweden, ICTMS2017-106-3.

Schmelzle S, Heethoff M, ... van de Kamp T (2017) The NOVA project: maximizing beam time efficiency through synergistic analyses of SRμCT data", Proc. SPIE 10391, Developments in X-Ray Tomography XI, 103910P-1.

Dormann CF, Blüthgen N (2017) Food webs versus interaction networks: principles, pitfalls and perspectives. In: Moore JC, de Ruiter PC, McCann KS, Wolters V (eds) Adaptive food webs: stability and transitions of real and model ecosystems, Cambridge University Press.

Frank K, Brückner A, Hilpert A, Heethoff M, Blüthgen N (2017) Nutrient quality of vertebrate dung as a diet for dung beetles. Scientific Reports 7: 12141

Mangels J, Fiedler K, Schneider FD, Blüthgen N (2017) Diversity and trait composition of moths respond to land-use intensification in grasslands: generalists replace specialists. Biodiversity and Conservation 26: 3385-3405

Staab M, Fornoff F, Klein AM, Blüthgen N (2017) Ants at plant wounds – a little-known trophic interaction with evolutionary implications for ant-plant interactions. American Naturalist 190: 442-450

Brückner A, Klompen H, Bruce AI, Hashim R, von Beeren C (2017). Infection of army ant pupae by two new parasitoid mite species (Mesostigmata: Uropodina). PeerJ 5: e3870

Brückner A, Hilpert A, Heethoff M (2017). Biomarker function and nutritional stoichiometry of neutral lipid fatty acids and amino acids in oribatid mites. Soil Biology & Biochemistry 115: 35-43.

Rosumek, FB, Brückner A, Blüthgen N, Menzel F, Heethoff M (2017). Patterns and dynamics of neutral lipid fatty acids in ants - implications for ecological studies. Frontiers in Zoology 14:36.

Tishechkin AKT, Kronauer DJC, von Beeren C (2017). Taxonomic review and natural history notes of the army ant-associated beetle genus Ecclisister Reichensperger (Histeridae: Haeteriinae). Coleopterists Bulletin, 71: 279-288

Mody K, Collatz J, Bucharova A, Dorn S (2017) Crop cultivar affects performance of herbivore enemies and may trigger enhanced pest control by coaction of different parasitoid species. Agriculture, Ecosystems and Environment 245: 74-82

Eisenhauer N, ... , Heethoff M, ... Powell JR (2017) Priorities for research in soil ecology. Pedobiologia 63: 1-7

Knuff A, Obermaier E, Mody K (2017) Differential susceptibility and suitability of domestic and wild apple species for a florivorous weevil and its parasitoids. Journal of Applied Entomology 141: 285-299

Brückner A, Heethoff M (2017) The ontogeny of oil gland chemistry in the oribatid mite Archegozetes longisetosus Aoki (Oribatida, Trhypochthoniidae). International Journal of Acarology 43: 337-342


Frank K, Hülsmann M, Assmann T, Schmitt T, Blüthgen N (2017) Land use affects dung beetle communities and their ecosystem services in forests and grasslands. Agriculture, Ecosystems and Environment 243: 114-122

Brückner A, Heethoff M, Blüthgen N (2017) The relationship between epicuticular long-chained hydrocarbons and surface area – volume ratios in insects (Diptera, Hymenoptera, Lepidoptera). PLoS One 12: e0175001

Kaluza BF, Wallace H, Keller A, Heard TA, Jeffers B, Drescher N, Blüthgen N, Leonhardt SD (2017) Generalist social bees maximize diversity intake in plant species-rich and resource-abundant environments. Ecosphere 8: e01758

Brückner A, Raspotnig G, Wehner K, Meusinger R, Norton RA, Heethoff M (2017) Storage and release of hydrogen cyanide in a chelicerate (Oribatula tibialis). Proceeding of the National Academy of Sciences USA, 114(13): 3469-3472

Koerner L, Garamszegi LZ, Heethoff M, Betz O (2017) Divergent morphologies of adhesive predatory mouthparts of Stenus species (Coleoptera: Staphylinidae) explain differences in adhesive performance and resulting prey-capture success. Zoological Journal of the Linnean Society, DOI: 10.1093/zoolinnean/zlx006

​​von Beeren C, Tishechkin AKT (2017). Nymphister kronaueri von Beeren & Tishechkin sp. nov., an army ant-associated beetle species (Coleoptera: Histeridae: Haeteriinae) with an exceptional mechanism of phoresy. BMC Zoology, 2:3 DOI: 10.1186/s40850-016-0010-x.


Kaiser-Bunbury CN, Mougal J, Whittington AE, Valentin T, Gabriel R, Olesen JM, Blüthgen N (2017) Ecosystem restoration strengthens pollination networks. Nature 542: 223-227

Brückner A, Heethoff M (2017) A chemo-ecologists’ practical guide to compositional data analysis. Chemoecology 27: 33-46 (with Erratum: Chemoecology 27: 47).

Wurmitzer C, Blüthgen N, Krell FT, Maldonado B, Ocampo F, Müller JK, Schmitt T (2017) Attraction of dung beetles to herbivore dung and synthetic compounds in a comparative field study. Chemoecology 27: 75-84


Kämper W, Blüthgen N, Eltz T (2017) Bumblebee footprints on bird’s-foot trefoil uncover increasing flower visitation with land-use intensity. Agriculture, Ecosystems and Environment 240: 77–83

Blüthgen N, Kaiser-Bunbury CN, Junker RR (2017) Biological invasions and ant-flower networks on islands. In: Oliveira PS, Koptur S. Ant-Plant Interactions: Impacts of Humans on Terrestrial Ecosystems. Cambridge University Press


Simons N, Lewinsohn T, Blüthgen N, Buscot F, Boch S, Daniel R, Gossner MM, Jung K, Kaiser K, Müller J, Prati D, Renner SC, Socher S, Sonnemann I, Weiner CN, Werner M, Wubet T, Wurst S, Weisser WW (2017) Contrasting effects of grassland management modes on species-abundance distributions of multiple groups. Agriculture, Ecosystems and Environment 237: 143-153


Staab M, Methorst J, Peters J, Blüthgen N, Klein AM (2017) Tree diversity and nectar composition affect arthropod visitors on extrafloral nectaries in a diversity experiment. Journal of Plant Ecology 10: 201-221

Batáry P, Gallé R, Riesch F, Fischer C, Dormann CF, Mußhoff O, Császár P, Fusaro S, Gayer C, Happe A-K, Kurucz K, Molnár D, Rösch V, Wietzke A & Tscharntke T (2017) The former iron curtain still drives biodiversity-profit trade-offs in German agriculture. Nature Ecology & Evolution 1: 1279-1284

Rousselin A, Bevacqua D, Sauge MH, Lescourret F, Mody K, Jordan MO (2017) Harnessing the aphid life cycle to reduce insecticide reliance in apple and peach orchards. A review. Agronomy for Sustainable Development 37: 38-49


Schleuning M, Fründ J, Schweiger O, Welk E, Albrecht J, Albrecht M, Beil M, Benadi G, Blüthgen N, Bruelheide H, Böhning-Gaese K, Dehling M, Dormann CF, Exeler N, Farwig N, Harpke A, Hickler T, Kratochwil A, Kuhlmann M, Kuhn I, Michez D, Mudri-Stojnic S, Plein M, Rasmont P, Schwabe A, Settele J, Vujic A, Weiner C, Wiemers M, Hof C (2016) Ecological networks are more sensitive to plant than to animal extinction under climate change. Nature Communications 7: 13965

Gossner MM, Lewinsohn T, Kahl T, Grassein F, Boch S, Prati D, Birkhofer K, Renner SC, Sikorski J, Wubet T, Arndt H, Baumgartner V, Blaser S, Blüthgen N, ... Weiner CN, Werner M, Wolters V, Wurst S, Westphal C, Fischer M, Weisser WW, Allan E (2016) Land-use intensification causes multitrophic homogenisation of grassland communities. Nature 540: 266-269

Kämper W, Weiner C, Kühsel S, Storm C, Eltz T, Blüthgen N (2016) Evaluating the effects of floral resource specialisation and of nitrogen regulation on the vulnerability of social bees in agricultural landscapes? Apidologie DOI 10.1007/s13592-016-0480-4

Bucharova A, Frenzel M, Mody K, Parepa M, Durka W, Bossdorf O (2016) Plant ecotype affects interacting organisms across multiple trophic levels. Basic and Applied Ecology 17: 688-695

Kriesell L, Hilpert A, Leonhardt SD (2016) Different but the same: bumblebee species collect pollen of different plant sources but similar amino acid profiles. Apidologie DOI 10.1007/s13592-016-0454-6

Kämper W, Werner PK, Hilpert A, Westphal C, Blüthgen N, Eltz T, Leonhardt SD (2016) How landscape, pollen intake and pollen quality. Landscape Ecology 31: 2245-2258


von Beeren C, Maruyama, M, and Kronauer DJC (2016). Community sampling and integrative taxonomy reveal new species and host specificity in the army ant-associated beetle genus Tetradonia (Coleoptera, Staphylinidae, Aleocharinae) PLoS ONE 11(11): e0165056.


Chisté MN, Mody K, Gossner MM, Simons NK, Köhler G, Weisser WW, Blüthgen N (2016) Losers, winners, and opportunists: how grassland land-use intensity affects orthopteran communities. Ecosphere 7: e01545

Heethoff M, Norton RA, Raspotnig G (2016) Once again: oribatid mites and skin alkaloids in poison frogs. Journal of Chemical Ecology 42: 841-844.

Brückner A, Wehner K, Neis M, Heethoff M (2016) Attack and defense in a gamasid-oribatid mite predator-prey experiment – sclerotization outperforms chemical repellency. Acarologia 56: 451-461

Soliveres S, van der Plas F, Manning P, Prati D, Gossner MM, Renner SC, Alt F, Arndt H, Baumgartner V, Binkenstein J, Birkhofer K, Blaser S, Blüthgen N, Boch S, ... Weiner CN, Weisser WW, Werner M, Westphal C, Wilcke W, Wolters V, Wubet T, Wurst S, Fischer M, Allan E (2016) Biodiversity at multiple trophic levels is needed for ecosystem multifunctionality. Nature 536: 456-459

Esser HJ, Herre EA, Blüthgen N, Loaiza JR, Bermúdez S, Jansen PA (2016) Host specificity in a diverse Neotropical tick community: an assessment using quantitative network analysis and host phylogeny. Parasites and Vectors 9: 372

Ellwood MDF, Blüthgen N, Fayle TM, Foster WA, Menzel F (2016) How do pairwise interactions affect community structure of epiphyte-dwelling ants in a tropical rainforest? Acta Oecologia 75: 24-34

Heethoff M, Scheu S (2016) Reliability of isotopic fractionation (15N, 13C) for the delimitation of trophic levels of oribatid mites: Diet strongly affects D13C but not D15N. Soil Biology and Biochemistry 101: 124-129

Kühsel S, Brückner A, Schmelzle S, Heethoff M, Blüthgen N (2016) Surface area – volume ratios in insects. Insect Science DOI 10.1111/1744-7917.12362

Wehner K, Norton RA, Blüthgen N, Heethoff M (2016) Specialization of oribatid mites to forest microhabitats – the enigmatic role of litter. Ecosphere 7: e01336

Sendoya S, Blüthgen N, Tamashiro J, Fernandez F, Oliveira P (2016) Foliage-dwelling ants in a neotropical savanna: effects of plant and insect exudates on ant communities. Arthropod-Plant Interactions 10: 183-195

Soliveres S, Manning P, Prate D, Gossner MM, ... Blüthgen N, ... Fischer M, Allan E (2016) Locally rare species influence grassland ecosystem multifunctionality. Philosophical Transactions Biological Sciences 371: 20150269

Ivens ABF, von Beeren C, Blüthgen N, Kronauer DJC (2016) Studying the complex communities of ants and their symbionts using ecological network analysis. Annual Review of Entomology 61:353–71

Blüthgen N, Simons NK, Jung K, Prati D, Renner SC, Boch S, Fischer M, Hölzel N, Klaus VH, Kleinebecker T, Tschapka M, Weisser WW, Gossner MM (2016) Land use imperils plant and animal community stability through changes in asynchrony rather than diversity. Nature Communications 7:10697


Houadria M, Blüthgen N, Salas-Lopez A, Schmitt MI, Arndt J, Schneider E, Orivel J, Menzel F (2016) The relation between circadian asynchrony, functional redundancy and trophic performance in tropical ant communities. Ecology 97: 225–235


Brückner A, Heethoff M (2016) Scent of a mite – Origin and chemical characterization of the lemonlike flavor of mite-ripened cheeses. Experimental and Applied Acarology 69: 249-261


Hartmann K, Laumann M, Bergmann P, Heethoff M, Schmelzle S (2016) Development of the synganglion and morphology of the adult nervous system in the mite Archegozetes longisetosus Aoki (Chelicerata, Actinotrichida, Oribatida). Journal of Morphology 277: 537-548





Allan E, Manning P, ... Blüthgen N, ... Fischer M (2015) Land use intensification alters ecosystem multifunctionality via loss of biodiversity and changes to functional composition. Ecology Letters 18: 834–843


Arnan X, Blüthgen N, Molowny-Horas R, Retana J (2015) Thermal characterization of European ant communities along thermal gradients and its implications for community resilience to temperature variability. Frontiers in Ecology and Evolution 3: 138


Arnan X, Blüthgen N (2015) Using eco-physiological traits to predict climate and activity niches: lethal temperature and water loss in Mediterranean ants. Global Ecology and Biogeography 24: 1454–1464

Gámez-Virués S, Perović DJ, Gossner MM, Börschig C, Blüthgen N, ... Tscharntke T, Westphal C (2015) Landscape simplification filters species traits and drives biotic homogenization. Nature Communications 6: 8568


Heethoff M, Rall BC (2015) Reducible defence: chemical protection alters the dynamics of predator-prey interactions. Chemoecology 25: 53-61


Houadria M, Salas-Lopez A, Orivel J, Blüthgen N, Menzel F (2015) Dietary and temporal niche differentiation in species-rich assemblages – can they explain local tropical ant coexistence? Biotropica 47: 208–217


Junker RR, Blüthgen N, Keller A (2015) Functional and phylogenetic diversities of plant communities differently affect the structure of flower-visitor interactions and reveal convergences in floral traits. Evolutionary Ecology 29: 437–450


Kaiser-Bunbury C, Blüthgen N (2015) Integrating network ecology with applied conservation: a synthesis and guide to implementation. AoB Plants 7: plv076


Kaiser-Bunbury C, Mougal J, Valentin T, Gabriel R, Blüthgen N (2015) Herbicide application as habitat restoration tool: impact on native plant communities. Applied Vegetation Science 18: 650–660


Kühsel S, Blüthgen N (2015) High diversity stabilizes the thermal resilience of pollinator communities in intensively managed grasslands. Nature Communications 6: 7989


Leonhardt SD, Wallace H, Blüthgen N, Wenzel F (2015) Potential role of environmentally derived cuticular compounds in stingless bees. Chemoecology 25: 159–167


Maglianesi MA, Blüthgen N, Böhning–Gaese K, Schleuning M (2015) Functional structure and specialization in three tropical plant–hummingbird interaction networks across an elevational gradient in Costa Rica. Ecography 38: 1119–1128


Mangels J, Blüthgen N, Frank K, Grassein F, Hilpert A, Mody K (2015) Tree species composition and harvest intensity affect herbivore density and leaf damage on beech, Fagus sylvatica, in different landscape contexts. PLoS One 10: e0126140


Manning P, Gossner MM, Bossdorf O, Allan E, Zhang YY, Prati D, Blüthgen N, ... Weisser WW, Fischer M (2015) Grassland management intensification weakens the associations among the diversities of multiple plant and animal taxa. Ecology 96: 1492–1501

Mody K, Collatz J, Dorn S (2015) Plant genotype and the preference and performance of herbivores: cultivar affects apple resistance to the florivorous weevil Anthonomus pomorum. Agricultural and Forest Entomology 17: 337-346

Renoult JP, Blüthgen N, Binkenstein J, Weiner CN, Werner M, Schaefer HM (2015) The relative importance of color signaling for plant generalization in pollination networks. Oikos 124: 347–354


Schmelzle S, Norton RA, Heethoff M (2015) Mechanics of the ptychoid defense mechanism in Ptyctima (Acari, Oribatida): One problem, two solutions. Zoologischer Anzeiger 254: 27-40


Staab M, Blüthgen N, Klein AM (2015) Tree diversity alters the structure of a tri-trophic network in a biodiversity experiment. Oikos 124: 827-834




Allan E, Bossdorf O, Dormann CF, Prati D, Gossner M, Tscharntke T, Blüthgen N, ... Weisser WW, Fischer M (2014) Inter-annual variation in land-use intensity enhances grassland multidiversity. Proceedings of the National Academy of Sciences USA 111: 308–313


Benadi G, Hovestadt T, Poethke HJ, Blüthgen N (2014) Specialization and phenological synchrony of plants and pollinators along an altitudinal gradient. Journal of Animal Ecology 83: 639–650


Maglianesi MA, Blüthgen N, Böhning–Gaese K, Schleuning M (2014) Morphological traits determine specialization and resource use in plant–hummingbird networks in the Neotropics. Ecology 95: 3325–3334

Riedel J, Dorn S, Mody K (2014) Assemblage composition of ants (Hymenoptera: Formicidae) affected by tree diversity and density in native timber tree plantations on former tropical pasture. Myrmecological News 20: 113-127

Weiner CN, Werner M, Linsenmair KE, Blüthgen N (2014) Land use impacts on mutualistic networks: disproportional declines in specialized pollinators via changes in flower composition. Ecology 95: 466–474




Albrecht J, Berens DG, Blüthgen N, Jaroszewicz B, Selva N, Farwig N (2013) Logging and forest edges reduce redundancy in plant–frugivore networks in an old-growth European forest. Journal of Ecology 101: 990–999


Benadi G, Blüthgen N, Hovestadt T, Poethke HJ (2013) When can plant-pollinator interactions promote plant diversity? American Naturalist 182: 131–146


Benadi G, Blüthgen N, Hovestadt T, Poethke HJ (2013) Contrasting specialization–stability relationships in plant–animal mutualistic systems. Ecological Modelling 258: 65–73


Bohan DA, Raybould A, Mulder C, Woodward G, Tamaddoni-Nezhad A, Blüthgen N, Pocock MJO, Muggleton S, Evans DM, Astegiano J, Massol F, Loeuille N, Petit S, Macfadyen S (2013). Networking agroecology: integrating the diversity of agroecosystem interactions. In: Woodward G, Bohan DA (eds.) Ecological Networks in an Agricultural World. Advances in Ecological Research 49: 1–67


Chrobock T, Weiner CN, Werner M, Blüthgen N, Fischer M, van Kleunen M (2013) Effects of native pollinator specialization, self-compatibility and flowering duration of European plant species on their invasiveness elsewhere. Journal of Ecology 101: 916–923


Heethoff M, Bergmann P, Laumann M, Norton RA (2013) The 20th anniversary of a model mite: A review of current knowledge about Archegozetes longisetosus (Acari, Oribatida). Acarologia 53: 353-368


Junker RR, Blüthgen N, Brehm T, Binkenstein J, Paulus J, Schaefer M, Stang M (2013) Specialisation on traits as basis for the niche-breadth of flower visitors and as structuring mechanism of ecological networks. Functional Ecology 27: 329–341


Heethoff M, Bergmann P, Laumann M, Norton RA (2013) The 20th anniversary of a model mite: A review of current knowledge about Archegozetes longisetosus (Acari, Oribatida). Acarologia 53: 353-368


Menzel F, Blüthgen N, Tolasch T, Conrad J, Beifuß U, Beuerle T, Schmitt T (2013) Crematoenones - a novel substance class exhibited by ants functions as appeasement signal. Frontiers in Zoology 10: 32

Riedel J, Dorn S, Brand G, Barrios H, Mody K (2013) Effects of ants on arthropod assemblages of a native timber tree in a tropical reforestation plantation. Journal of Applied Entomology 137: 418-428

Riedel J, Dorn S, Plath M, Potvin C, Mody K (2013) Time matters: temporally changing effects of planting schemes and insecticide treatment on native timber tree performance on former pasture. Forest Ecology and Management 297: 49-56

Riedel J, Dorn S, Plath M, Mody K (2013) Growth, herbivore distribution, and herbivore damage of timber trees in a tropical silvopastoral reforestation system. Annals of Forest Science 70: 75-86




Benadi G, Blüthgen N, Hovestadt T, Poethke HJ (2012) Population dynamics of interacting plant and pollinator communities: stability reconsidered. American Naturalist 179: 157–168


Blüthgen N, Dormann CF, Prati D, Klaus VH, Kleinebecker T, Hölzel N, Alt F, Boch S, Gockel S, Hemp A, Müller J, Nieschulze J, Renner SC, Schöning I, Schumacher U, Socher SA, Wells K, Birkhofer K, Buscot F, Oelmann Y, Rothenwöhrer C, Scherber C, Tscharntke T, Weiner CN, Fischer M, Kalko EKV, Linsenmair KE, Schulze ED, Weisser WW (2012) A quantitative index of land-use intensity in grasslands: integrating mowing, grazing and fertilization. Basic and Applied Ecology 13: 207–220


Leonhardt SD, Blüthgen N (2012) The same, but different: pollen foraging in honeybee and bumblebee colonies. Apidologie 43: 449-464


Menzel F, Staab M, Chung AYC, Gebauer G, Blüthgen, N (2012) Trophic ecology of parabiotic ants: Do the partners have similar food niches? Austral Ecology 37: 537–546


Polidori C, Santoro D, Blüthgen N (2012) Does prey mobility affect niche width and individual specialization in hunting wasps? A network-based analysis. Oikos 122: 385–394


Schleuning M, Fründ J, Klein AM, Abrahamczyk S, Alarcón R, Albrecht M, Andersson GKS, Bazarian S, Böhning-Gaese K, Bommarco R, Dalsgaard B, Dehling DM, Gotlieb A, Hagen M, Hickler T, Holzschuh A, Kaiser-Bunbury CN, Kreft H, Morris RJ, Sandel B, Sutherland WJ, Svenning JC, Tscharntke T, Watts S, Weiner CN, Werner M, Williams NM, Winqvist C, Dormann CF, Blüthgen N (2012) Specialization of mutualistic interaction networks decreases towards tropical latitudes. Current Biology 22: 1925–1931




Bleil R, Blüthgen N, Junker RR (2011) Ant-plant mutualism in Hawai’i? Invasive ants reduce flower parasitism but also exploit floral nectar of the endemic shrub Vaccinium reticulatum (Ericaceae). Pacific Science 65: 291–300


Blüthgen N, Klein AM (2011) Functional complementarity and specialisation: The role of biodiversity in plant-pollinator interactions. Basic and Applied Ecology 12: 282–291


Drescher J, Feldhaar H, Blüthgen N (2011) Interspecific aggression, resource monopolization and ecological dominance of Anoplolepis gracilipes within an ant community in Malaysian Borneo. Biotropica 43: 93–99


Junker RR, Bretscher S, Dötterl S, Blüthgen N (2011) Phytochemical cues affect hunting site choices of a nursery web spider (Pisaura mirabilis) but not of a crab spider (Misumena vatia). Journal of Arachnology 39: 113–117


Junker RR, Daehler CC, Dötterl S, Keller A, Blüthgen N (2011) Hawaiian ant–flower networks: nectar-thieving ants prefer undefended native over introduced plants with floral defenses. Ecological Monographs 81: 295–311


Klaus VH, Kleinebecker T, Hölzel N, Blüthgen N, Boch S, Müller J, Socher S, Prati D, Fischer M (2011) Nutrient concentrations and fibre contents of plant community biomass reflect species richness patterns along a broad range of land-use intensities in agricultural grasslands. Perspectives in Plant Ecology, Evolution and Systematics 13: 287–295


Leonhardt SD, Form S, Blüthgen N, Schmitt T, Feldhaar H (2011) Genetic relatedness and chemical profiles in an unusually peaceful eusocial bee. Journal of Chemical Ecology 37: 1117–1126


Leonhardt SD, Blüthgen N & Schmitt T (2011) Chemical profiles of body surfaces and nests from six Bornean stingless bee species. Journal of Chemical Ecology 37: 98–104


Leonhardt SD, Schmitt T, Blüthgen N (2011) Tree resin composition, collection behavior and selective filters shape chemical profiles of tropical bees (Apidae: Meliponini). PLoS One 6: e23445


Junker R, Loewel C, Gross R, Dötterl S, Keller A, Blüthgen N (2011) Composition of epiphytic bacterial communities differs on petals and leaves. Plant Biology 13: 918–924


Schleuning M, Blüthgen N, Flörchinger M, Braun J, Schaefer HM, Böhning-Gaese K (2011) Specialization and interaction strength in a tropical seed-dispersal network differ among forest strata. Ecology 92: 26–36


Weiner CN, Werner M, Linsenmair KE, Blüthgen N (2011) Land use intensity in grasslands: changes in biodiversity, species composition and specialisation in flower-visitor networks. Basic and Applied Ecology 12: 292–299



(The lab group started in 2011 in Darmstadt; for publications before 2011, see individual people's pages)